One inclusion formed somewhat later exterior to the exosporium. Another inclusion was formed within the exosporium and remained with the spore after mother cell lysis.

当芽形解后，一种晶体被释放出来，形自由的伴晶体，另一种在芽内侧形，熟后被包裹在芽以内不脱落，始终和芽粘连在一起。